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owls are among the most visually cryptic birds, and some of their behaviours are quite unique. to date, the vocal repertoire of owls has been poorly studied (gonzález et al. 2001 ; ramos et al. 2003 ; ramos & gonzález 2003 ; ramos & walz 2005 ; ramos & lorenzoni 2005 ; ramos et al. 2006 ; eith & morrison 2009 ; eith et al. 2009 ; ramírez, muñoz & gonzález 2010 ; ramírez, muñoz, álvarez & gonzález 2014 ). most of the vocalizations of owls can be divided into two groups: long series of calls and screams. the first group is composed of two or more repeated calls (long calls), that are short in duration and repeated at intervals shorter than 1 s (ramos et al. 2003 ). studies of different individual owls have demonstrated that the vocal repertoire of each species of owls presents a large degree of individual variation (ramos et al. 2003 ; ramos & gonzález 2003 ; ramos & lorenzoni 2005 ; ramos et al. 2006 ; eith et al. 2009 ). thus, there is a wide variation in the spectral content and duration of the long calls between individuals and even within the same individual during the breeding season (ramos, muñoz & gonzález 2010 ). the second group is composed of a repeated scream that is used for protection, food calling, attracting conspecifics or as a threat. screams usually last 1-5 s and tend to have a large overlap in the frequency spectrum; they are produced by the whole body, with the tongue and larynx (martinez-demuynck, bell & marin 2001 ; gasparini 2003 ; norambuena & muñoz-pedreros 2012 ; ramírez et al. 2014 ).

owls can be seen in daylight during the day when they arrive from breeding grounds to feed in the early morning or after sunset (martín et al. 2004 ; gonzález et al. 2005a ; gonzález et al. 2005b ). ibarra et al. (2004) state that the long calls can be heard up to 3 km from a given individual. however, other authors have suggested that the detection radius for the long calls decreases as the number of calls increases (spear, hester & spear 2009 ; ramírez, muñoz & gonzález 2010 ). the long calls may be heard as a low-frequency signal; the short calls tend to have higher frequencies and may be more easily detected, depending on the area in which the survey is performed (spear et al. 2009 ). the long calls of owls can have a degree of frequency overlap and they can be heard simultaneously (ramos et al. 2006 ; ramos et al. 2010 ). the latter property can be used to obtain information about both identity and/or behavioural state of the calling individual (spear, hester & spear 2009 ; spear et al.

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